The current mill at Wellesbourne is the last of a long line of mills on the river Dene, dating back before the Doomsday. There were 3 mills in the manor of Walton in 1086, including the present one.

Little is known of the mills early history but From 1845 the Bustin family were tennant millers untill 1872. with the Anderson family following from 1876 to 1908.

T he mill last worked under water power at this time but carried on by external pulley driven from a tractor up to 1958 and was last used by Mr Bartlett for pig feed.

The mill was initially rebuilt/ restored in 1989-90 by the David Nicholls and Chilton Partnership (millwrights) and Sir. Andrew Hamilton the owner of the Walton estate.

The present three story building was put up in 1834 (a date exists in the south west gable) and has an internal breast shot wheel, of unusual design, having three sets of three pairs of wooden clasp arms. The wheel is 17ft diameter and 6 ft wide and generates around 20hp.

It drives two pairs of 4 ft diameter millstones (one peak for animal feed, and one burr pair for flour) from below in the usual manner, from a 7ft 4ins diameter wooden spur to a 18 inch stone nut on the stone spindle.